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Kadwell, M., Fernandez, M., Stanley, H.F., Baldi, R., Wheeler, J.C., Rosadio, R. and Bruford, M.W. (2001) Genetic analysis reveals the wild ancestors of the llama and the alpaca. Proceedings of the Royal Society B: Biological Sciences, 268, 2575-2584. doi:10.1098/rspb.2001.1774

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