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Stavenga, D.G., Leertouwer, H.L. and Wilts, B.D. (2014) Coloration Principles of Nymphaline Butterflies—Thin Films, Melanin, Ommochromes and Wing Scale Stacking. Journal of Experimental Biology, 217, 2171-2180.
http://dx.doi.org/10.1242/jeb.098673

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