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Agnese, J.F. and Teugels, G.G. (2001) Genetic Evidence for Monophyly of the Genus Heterobranchus and Paraphyly of the Genus Clarias (Siluriformes, Clariidae). Copeia, 2, 548-552.
http://dx.doi.org/10.1643/0045-8511(2001)001[0548:GEFMOT]2.0.CO;2

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